Interesting - I have read the whole article. Its a standard article using the common terms for the field. A few things:
1. The vapor was tested on mouse cells - further research is recommended by the authors.
2. Actual vapor, not liquid, was used. Important as previous research indicated e-liquid in its liquid state can cause cytotoxicity (poison cells). However in this study vapor was made using an ecig and atomizer - and the vapor was tested on the cells.
3. Cigarette smoke was also used.
4. Cigarette smoke was very cytotoxic. E-liquid vapor was not. The e-liquid used was close to 50/50 PG/VG. Only one flavor showed cytoxicity and that was a coffee flavor at a high concentration of coffee flavor, and still well less than cytotoxicity from cigarette smoke.
Just a simplified summary. The good thing about this, in my view, is that they replicated a vaping experience by using a 3.7v battery, an atomizer, and vapor. This is a really quite an effective demonstration of the difference between vapor and cigarette smoke.